Data S1. Text file of the alignment used for the phylogenetic analysis shown in Figure 3a.

>Zm00001d017427|Zea mays （Sh4/ZmYSL2）
MNDFRKLFSFNSTRLQMPRTTTPQGLLIYNGLVESEALIDSSSCSSRTAMEINPADEIDKCEAGGGAEPQPADPEPDAAA
AVEQQQPWREQLTARGLVAAALIGSMYTVVVMKLNLTTGFVPTMNVSAALLAFLALRGWTGALSRLGVAGARPFTRQENT
VVQTCAVACYSLALCGFGSFMLGLNRRTYEAAGVNTPGNVPGSVKEPGFGWIAGFLVASSFGGLLTLIPLRKALVIDYKL
TYPSGTATAVLINGFHTPQGDKNAEKQVRAFLKYFGISFLWSFFQWFYTGGNACGFAQFPTFGLKAWKHSFFFDFSMTYV
GAGMICPHLVNLSALLGAVLSWGILWPLISRRKGYWYPADVPESSMTSLYGYKSFVCIALIMGDGIYHLVKVLGVTAKSL
HERSKLRRSNNRGSSSRVADEDNAAAIDSLRRDEVFNRDSIPAWLAYAGYAVTSAVAAVAIPAMFRQVRWYYVVAAYALA
PVLGFSNSYGAGLTDINMGYSYGQLALFVLAAWAGRDDGVVAGLVGCGLVKQLVLISADLMHDFKTAHLTLTSPRSMLAA
QAVGAAMGCVVTPLTFLLFYRAFDVGNPDGYWKAPFALIYRNMALLGAQGFSALPTHCLPLSAGFFALAVLANVMKDVLP
PRYARYVPLPTAMAVPFLVGASFAIDMVVGTVVVFAWHWIDGNETTLLVPAVASGLICGDGVWTFPSSLLSLAKIKPPIC
LKFTPGS

>Zm00001d017429|Zea mays (YS1)
MDLARSGGAAAADDGGEIERHEPAPEDMESDPAAAREKELELERVQPWREQVTLRGVVAALLIGFMYSVIVMKIALTTGL
VPTLNVSAALMAFLALRGWTRVLERLGVAHRPFTRQENCVIETCAVACYTIAFGGGFGSTLLGLDKKTYELAGASPANVP
GSYKDPGFGWMAGFVAAISFAGLLSLIPLRKVLVIDYKLTYPSGTATAVLINGFHTKQGDKNARMQVRGFLKYFGLSFVW
SFFQWFYTGGEVCGFVQFPTFGLKAWKQTFFFDFSLTYVGAGMICSHLVNISTLLGAILSWGILWPLISKQKGEWYPANI
PESSMKSLYGYKAFLCIALIMGDGTYHFFKVFGVTVKSLHQRLSRKRATNRVANGGDEMAALDDLQRDEIFSDGSFPAWA
AYAGYAALTVVSAVIIPHMFRQVKWYYVIVAYVLAPLLGFANSYGTGLTDINMAYNYGKIALFIFAAWAGRDNGVIAGLA
GGTLVKQLVMASADLMHDFKTGHLTMTSPRSLLVAQFIGTAMGCVVAPLTFLLFYNAFDIGNPTGYWKAPYGLIYRNMAI
LGVEGFSVLPRHCLALSAGFFAFAFVFSVARDVLPRKYARFVPLPMAMAVPFLVGGSFAIDMCVGSLAVFVWEKVNRKEA
VFMVPAVASGLICGDGIWTFPSSILALAKIKPPICMKFTPGS

>XP_021315728.1|Sorghum bicolor
MARRQSPGGWGSPPLITPVHSSSCSSRSANMEINPADEIDKCEAGGGAEPQPADPEPEAASLELEQPQPW
REQLTARGLVAAALIGSMYTVVVMKLNLTTGFVPTMNVSAALLAFLALRGWTGALARLGAAGAGRPFTRQ
ENTVVTTCAVACYSLALCGFGSFILGLNKRTYEAAGVDTPGNVPGSVKEPGFGWITGFLVASSFGGLLTL
IPLRKALVIDYKLTYPSGTATAVLINGFHTPQGDKNAEKQVRAFLKYFGISFFWSFFQWFYTGGNACGFA
QFPTFGLKAWKQSFFFDFSMTYVGAGMICPHHVNLSTLLGAVLSWGILWPLIHNRKGDWYSADIPESSMT
SLYGYKSFVCIALIMGDGIYHFVKVLGVTAKSLHEQSKLRRGSNRVADEGNAAAIDDLRRDEVFNRDSIP
AWLAYAGYAVVSAVAVVAIPMMFRQVRWYYVVAAYALAPVLGFSNSYGAGLTDINMGFSYGQLALFVLAA
WAGRDDGVVAGLVGCGLVKQLVLISADLMHDFKTAHLTLTSPRSMLAAQAVGAAIGCVVTPLTFLLFYRA
FDVGNPNGYWKAPFALIYRNMALLGAQGFSALPTHCLPLSAGFFAFAVLANVVKDVLPPRYKRYVPLPTA
MAVPFLVGASFAIDMCVGTLVVFAWQRVNGNDSTLLVPAVASGLICGDGVWTFPSSLLSLAKIKPPMCLK
FTPGS

>XP_015625410.1|Oryza sativa Japonica Group
MEAAAPEIERCDAGDVESDHDGAAAAAERVPPWREQVTARGMVAALLIGFVYTVIIMKLALTTGIIPTLN
VSAALLAFLALRGWTRAPALLLPGGGAASSSSRRRPFTRQENTVVQTCAVACYTMGFGGARPATTSCGFG
SSLLALNRKTYELAGVSTPGNSPGSYKEPGVGWMTGFLFAISFVGLLNLLPLRKALIIDYKLTYPSGTAT
AVLINGFHTPQGENSAKKQVRGFLNCFGISLLWSFFQWFYTGGESCGFLQFPTFGLKAWKQTFYFDFSLT
YVGAGMICSHLVNLSALFGAILSWGIMWPLISIQKGKWYPGNVPESSMTSLFGYKSFMCVALIMGDGLYH
FIKVTGITAKSLHERSNRRHAKKATDEDTFVIADMQRDEFFNKDYIPNWLAYAGYALLSIVAVIAIPIMF
QQVKWYYVVVAFVLAPVLGFSNAYGTGLTDMNMSYNYGKIALFIFAAWGGRDNGVIAGLVGCGIVKQLVQ
VSADLMHDFKTGHLTLTSPRSMLVGQAIGTAMGCIIAPLTFLLFYKAFDIGNPDGYWKAPYALIFRNMAI
LGVEGFSALPKHCLELSAGFFAFSVLINLMRDFLPRKYRDYVPLPTAMAVPFLVGANFAIDMCVGSLIVF
AWHKINSKESALLVPAVASGFICGDGIWMFPSSLLSLAKVKPPICMKFTPGS

>XP_016740678.1|Gossypium hirsutum
MNMEEMKEIERVERQDSEEMNNEQEELKRIAPWMKQITIRGLVASLLIGIIYSVIVMKLNLTTGLVPNLN
VSAALLAFVFIRSWTKLLQKAGIVSTPFTRQENTIIQTCAVACYSIAVGGGFGSYLLGLNRKTYDQAGIG
TEGNNPWSIKEPGIGWMVGFLFVSCFVGLLALVPLRKIMIIDYKLSYPSGTATSVLINGFHTPKGDKIAK
KQVHGFVNFFSLSFLWAFFQWFYAGGDKCGFAQFPTFGLKAWKNSFYFDFSMTYIGAGMICSHLVNLSLL
LGAVLSWGVMWPLIGGLKGEWFPATLPESSMKSLNGYKVFISIALILGDGLYNFLKILFLIARGIHTNVK
VRSLKTFSHDQKQQQIHLQRNELFVRENIPIWVACAGYTIFSILSIVVIPLMFPELKWYYIVVAYILAPS
LSFCNAYGAGLTDMNMAYNYGKVALFVLAAMSGKENGVVAGLVGCGLIKSIVSISSDLMHDFKTGHLTLT
SPRSMLVSQAIGTAIGCVVAPLTFFLFYKAFDVGNPDGEYKAPYALIYRNMAILGVQGFSALPHHCLQLC
YGFFAFAIAANLLRDFSPKNIGKWVPLPMAMAVPFLVGAYFAIDMCVGSLVVFGWYKLNGKKADLMVPAV
ASGLICGDGLWLLPSSILALFKVRPPICMSFFAST

>XP_003527996.2 |Glycine max
MNPMGTSTVSDEELKEIENLGREDIEEAPIVPEDVSRIAPWIRQITLRGLVASFLIGIIYSVIVMKLNLT
TGLVPNLNVSAALLGFVLIRAWTKVLAKAKIVSTPFTRQENTIIQTCAVACYSIAVGGGFGSYLLGLNRR
TYEQAGVGTEGNNPGSTKEPGIGWMTAFLFVTSFVGLLALVPIRKIMIIDYKLTYPSGTATAVLINGFHT
PKGDVMAKKQVHGFLKFFSASFLWAFFQWFYSGGDNCGFVQFPTFGLKAWKNSFYFDFSMTYVGAGMICS
HLVNLSLLLGAVISWGIMWPLIRGLKGEWFPASIPESSMKSLNGYKVFISIALILGDGLYNFVKVLYFTA
TNIHATVKRKNPETFSDNQKPLPLDDLRRNEVFARESIPIWLACTGYILFSIVSIIVIPLMFPQLKWYYV
VFAYLFAPSLGFCNAYGAGLTDMNMAYNYGKVALFVLSALAGKNDGVVAGLVGCGLIKSIVSISSDLMHD
FKTGHLTFTSPRSMLLSQAIGTAIGCVVAPLTFFLFYKAFDVGNPDGDYKAPYAIIYRNMAILGVEGFSA
LPHHCLQLCYGFFTFAIAANLVRDLGPKKIGKWIPLPMAMAVPFLVGGYFAIDMCMGSLVVFLWHTLNRN
EAGLMVPAVASGLICGDGLWILPSSILALLKIRPPICMSFLSASAS

>XP_016489802.1|Nicotiana tabacum
MSRVGVMREEEKEKSMEIQREEMEEVRDYSEEVKRIPPWTKQITVRGIVASVLIGIIYSVIVTKLNLTTG
LVPNLNVSAALLAYVFIQSWTKILKKANFVYTPFTRQENTIIQTCAVACYSIAVGGGFGSYLLGLNKKTY
EQAGVDTEGNTPGSHKEPSLDWMIGFLFVVSFVGLLALVPLRKIMIIDYKLAYPSGTATAVLINGFHTPK
GDKMAKKQVKGFMKVFSMSFLWSFFQWFYSGGDQCGFAQFPTFGLKAWKQSFYFDFSMTYVGAGMICSHL
VNLSLLLGAVLSWGIMWPLIGERKGSWFPATLPQSSMKSLSGYKVFISIALILGDGLYNFVRTLFFTGRS
IYVSLKTRRPESSAAADNKNQPLDELQRNEIFIRESIPLWLACIGYMVFSLISIIVIPLMFPALKWYYVL
VAYILAPALSFCNAYGAGLTDLNMAYNYGKVALFVLAALSGKENGVVAGLIGCGLIKSIVSISSDLMHDF
KTSHLTLTSPRSMLLSQAIGTAIGCVVAPLTFFLFYKAFPVGDPNGDYKAPYAIVYRNMAILGVEGFSAL
PNHCLQLCYGFFAFAIIANLVRDITPERFGKWVPLPMAMAVPFLNNASFAIDMCVGSLIVYVWHKLNSKK
ASLMVPAVASGLICGDGLWILPSALLALVKVKPPICMAFTVGQT

>NP_197826.2 |Arabidopsis thaliana
MENERVEREQSQFQEDEFIDSRKPPPWRKQITVRAIVASLLIGIVYSVICLKLNLTTGLVPNLNISSALL
AFVFLKSWTKVLQKAGIATTPFTRQENTIAQTCAVACYSISLAGGFASYLLGLNRRTYEETGVNTEGNNP
RGIKEPGVGWMTSFLFVTSFIGLVVLVPLRKVMIIDYKLTYPSGTATAVLINGFHTSKGDKTAKKQIRGF
IKSFGLSFFWAFFGWFYSGGEKCGFSQFPTFGLQALDKTFYFDFSMTYVGAGMICSHLVNLSLLFGAILS
WGIMWPLIARLKGEWFPATLKDNSMQGLNGYKVFICIALILGDGLYNFVKILFFTGRSFHSRLSKTNSIS
TLVEVPEDSTKESDNLKRENEVFVRESIPLWMACVGYLFFSLVSIIAIPLMFPQLKWYFVLVAYLLAPSL
SFCNAYGAGLTDMNMAYNYGKAALFVMAALAGKNDGVVAGMVACGLIKSIVSVSADLMHDFKTGHLTQTS
PRSMLVAQAIGTAIGCVVAPLTFFLFYKAFDVGNQNGEYKAPYAMIYRNMAIIGVQGPSALPKHCLELCY
GFFAFAVAANLARDLLPDKPGKWIPLPMAMAVPFLVGGSFAIDMCIGSLVVYVWKKVNRKKADVMVPAVA
SGLICGDGLWILPSSLLALAKVRPPICMNFTAAH

>XP_024380865.1 |Physcomitrium patens
MGRDNGKEAGPVRRSYHGAQNGSDAMDTTIGGNEMPGNDFKVSDMAVPKSHGVYNVNDDHVGLEDAFMLV
DIPRWQDQLTPRGLFVSFVLGTLFCIITHKLNLTVGVIPSLNISAGLLGFFFIRLWALVSSKLGFFTTPF
TRQENTVIQTCVVACYGLAFSGGFGSYILGMSQKTHDQVNTAAPGDIPGNEGVKNPGLGWMFAFIVVVSF
VGIFSIVILRKVMIIDYRLTYPSGTATGILINSFHTPGGVEVAKKQVRCLGKCFTVSFLWGFFKWFFSGI
GNSCGFDNFPTLGFKAYHNKMYFDFSMTYIGAGIISPHIVNVSVLLGAVVAWGLMWPLISNHEGDWYPTG
LKETDFRGLYSYKVFIAIAVILGDGLYNFLKIIFKSARQLYVHYKTNNQLPVTAPAKLDAMDRDHARRTE
IFMKEGIPQWVALLGYIFLAIVSTISMPHLFSPVKWYYVLICYSLAPILAFCNAYGTGLTDWNLSSSYGK
LALFVFGAWAGTDGGVLVGLALCGVMMSIVSVAADLIQDFKTGYLTLSSPRSMFVSQLVGTLMGCVLAPS
TFWLFYKAYSIGDPDGVYKAPYAVVYRSMALIGVEGFSALPSHCLEICYCLFVAAVLINLLRDFLPRRIG
QFVPIPMAMAIPFYIGGYFAIDMFIGSVIRFVYEKVNKAKADIITPAIAAGLICGDGVWTIPSSILALTK
VNPPICMYFYSSSNPLAGTSY